*'''Victus''' - Its execution techniques have been used by Champions to claim the lives of countless tyrants and warlords.{{Fn|41}}

The source of the [[gene-seed]] of the Custodes is a mysterious issue, if indeed they even ''have'' something similar enough to [[space marine]]s for the term "gene-seed" to have meaning. Some say that the Custodian Guard are to the Emperor what the Space Marines are to the Primarchs - that the Emperor's own genetic matrix was used in their creation and through this, their loyalty to him is assured.{{Fn|14a}} Others argue that the Custodians are not like the Emperor in the way that a Space Marine is like his Primarch, and that some other source was used as a template for their physical and psychological form, a source that was lost during the anarchy of the Age of Strife.{{Fn|12}} Recent sources, however, seem to firmly establish that the Custodes were created from the flesh and blood of the Emperor himself. {{Fn|19}} The genetic enhancement that forms the Custodes is different from and predates that developed to create the [[Space Marines]].{{Fn|3}} Each Custodian is a unique work of art, the product of genetic lore collected over many lifetimes{{Fn|15b}}.